:root{
  --c1:#233C4B; /* Blue Dianne */
  --c2:#FF7D2D; /* Crusta */
  --c3:#FAC846; /* Bright Sun */
  --c4:#A0C382; /* Olivine */
  --c5:#5F9B8C; /* Patina */

  /* Light theme (same palette, brighter surfaces) */
  --bg: #ffffff;
  --bg-soft: #f7faf9;
  --panel: #ffffff;
  --panel2: rgba(35,60,75,0.04);
  --panel3: rgba(35,60,75,0.06);
  --text: rgba(35,60,75,0.95);
  --muted: rgba(35,60,75,0.72);
  --border: rgba(35,60,75,0.14);

  --shadow: 0 16px 40px rgba(35,60,75,0.16);
  --shadow-soft: 0 12px 30px rgba(35,60,75,0.10);
  --radius: 18px;
  --radius-sm: 12px;

  --container: 1120px;
  --font: ui-sans-serif, system-ui, -apple-system, "Segoe UI", Roboto, Arial, "Helvetica Neue", Helvetica, sans-serif;
}

*{box-sizing:border-box}
html{scroll-behavior:smooth}
body{
  margin:0;
  font-family:var(--font);
  color:var(--text);
  background:
    radial-gradient(1000px 520px at 12% 0%, rgba(95,155,140,0.18), transparent 60%),
    radial-gradient(900px 520px at 88% 12%, rgba(250,200,70,0.16), transparent 60%),
    radial-gradient(1100px 700px at 55% 92%, rgba(160,195,130,0.14), transparent 60%),
    linear-gradient(180deg, var(--bg) 0%, var(--bg-soft) 100%);
}

a{color:inherit}
code{
  font-family: ui-monospace, SFMono-Regular, Menlo, Monaco, Consolas, "Liberation Mono", "Courier New", monospace;
  background: var(--panel3);
  padding: 0.15rem 0.35rem;
  border-radius: 8px;
  border: 1px solid var(--border);
}

.skip-link{
  position:absolute;
  top:-60px; left:16px;
  padding:10px 12px;
  background:var(--c3);
  color:#111;
  border-radius:10px;
  z-index:999;
  transition: top 0.2s ease;
}
.skip-link:focus{top:16px}

.container{
  width:min(var(--container), calc(100% - 40px));
  margin:0 auto;
}

.site-header{
  position:relative;
  overflow: visible;
}

.nav{
  display:flex;
  align-items:center;
  justify-content:space-between;
  gap:18px;
  padding:18px 0;
}

.brand{
  display:flex;
  align-items:center;
  gap:12px;
  text-decoration:none;
}
.brand-mark{
  width:40px; height:40px;
  display:grid; place-items:center;
  background: linear-gradient(135deg, var(--c2), var(--c3));
  color:#111;
  border-radius: 14px;
  font-weight: 800;
  box-shadow: var(--shadow-soft);
}
.brand-text{font-weight:700; letter-spacing:0.2px}

.nav-links{
  display:flex;
  align-items:center;
  gap:14px;
}
.nav-links a{
  text-decoration:none;
  padding:10px 10px;
  border-radius: 12px;
  color: var(--muted);
  border:1px solid transparent;
}
.nav-links a:hover{
  color:var(--text);
  border-color: var(--border);
  background: var(--panel2);
}
.nav-cta{
  background: rgba(160,195,130,0.22);
  border-color: rgba(160,195,130,0.45) !important;
  color: var(--text) !important;
}

.nav-toggle{
  display:none;
  background: transparent;
  border: 1px solid var(--border);
  color: var(--text);
  border-radius: 12px;
  padding:10px 12px;
  cursor:pointer;
}

.hero{
  position:relative;
  padding: 28px 0 56px;
}
.hero-bg{
  z-index: 0;
  position:absolute;
  inset:-40px -40px auto -40px;
  height: 340px;
  background:
    linear-gradient(90deg,
      rgba(35,60,75,0.22) 0%,
      rgba(255,125,45,0.22) 22%,
      rgba(250,200,70,0.22) 46%,
      rgba(160,195,130,0.22) 72%,
      rgba(95,155,140,0.22) 100%);
  filter: blur(26px);
  opacity:0.85;
}

.hero-inner{
  position:relative;
  z-index: 2;
  display:grid;
  grid-template-columns: 1.4fr 0.9fr;
  gap: 22px;
  align-items: start;
}

.hero-copy h1{
  margin: 0 0 10px 0;
  font-size: clamp(2.0rem, 3.0vw, 3.0rem);
  line-height: 1.05;
  letter-spacing: -0.02em;
}
.hero-title{
  margin: 0 0 14px 0;
  color: var(--muted);
  display:flex;
  flex-wrap:wrap;
  gap:10px;
  align-items:center;
}
.badge{
  display:inline-flex;
  align-items:center;
  padding: 6px 10px;
  border-radius: 999px;
  border: 1px solid rgba(250,200,70,0.35);
  background: rgba(250,200,70,0.12);
  color: var(--text);
  font-weight: 700;
  font-size: 0.92rem;
}

.hero-lead{
  margin: 0 0 18px 0;
  font-size: 1.05rem;
  line-height: 1.55;
  max-width: 62ch;
}

.hero-actions{
  display:flex;
  flex-wrap:wrap;
  gap:12px;
  margin-bottom: 16px;
}
.btn{
  display:inline-flex;
  align-items:center;
  justify-content:center;
  gap:10px;
  text-decoration:none;
  padding: 12px 16px;
  border-radius: 14px;
  border: 1px solid var(--border);
  box-shadow: var(--shadow-soft);
}
.btn.primary{
  background: linear-gradient(135deg, rgba(255,125,45,0.95), rgba(250,200,70,0.92));
  color:#141414;
  border-color: rgba(255,255,255,0.10);
  font-weight: 700;
}
.btn.ghost{
  background: var(--panel2);
  color: var(--text);
}
.btn:hover{transform: translateY(-1px)}
.btn:active{transform: translateY(0px)}

.hero-meta{
  display:flex;
  flex-wrap:wrap;
  gap:10px;
}
.chip{
  text-decoration:none;
  padding: 8px 10px;
  border-radius: 999px;
  border: 1px solid var(--border);
  background: rgba(255,255,255,0.05);
  color: var(--muted);
}
.chip:hover{color:var(--text); background: rgba(255,255,255,0.08)}

.hero-card{
  background: var(--panel2);
  border: 1px solid var(--border);
  border-radius: var(--radius);
  box-shadow: var(--shadow-soft);
  padding: 18px;
  backdrop-filter: blur(10px);
}
.card-title{margin:0 0 10px 0; font-size:1.1rem}
.card-text{margin:0 0 14px 0; color:var(--muted); line-height:1.55}

.stats{
  display:grid;
  gap:10px;
}
.stat{
  padding: 10px 12px;
  border-radius: 14px;
  border: 1px solid var(--border);
  background: rgba(0,0,0,0.10);
}
.stat-k{color: var(--muted); font-size: 0.9rem}
.stat-v{margin-top: 4px; font-weight: 600}

.mini-actions{margin-top: 10px}
.text-link{
  color: var(--c4);
  text-decoration:none;
  font-weight:700;
}
.text-link:hover{text-decoration:underline}

.section{
  padding: 64px 0;
}
.section.alt{
  background: rgba(35,60,75,0.03);
  border-top: 1px solid var(--border);
  border-bottom: 1px solid var(--border);
}
.section h2{
  margin: 0 0 10px 0;
  font-size: 1.8rem;
  letter-spacing: -0.01em;
}
.section-lead{
  margin: 0 0 22px 0;
  color: var(--muted);
  line-height: 1.65;
  max-width: 80ch;
}

.grid{
  display:grid;
  gap: 16px;
}
.cards-4{
  grid-template-columns: repeat(4, minmax(0,1fr));
}
.two-cols{
  grid-template-columns: repeat(2, minmax(0,1fr));
}

.card{
  background: var(--panel);
  border: 1px solid var(--border);
  border-radius: var(--radius);
  padding: 16px;
  box-shadow: var(--shadow-soft);
}
.card h3{margin: 4px 0 10px 0}
.card p{margin:0; color:var(--muted); line-height:1.6}
.icon{
  width: 44px; height: 44px;
  border-radius: 16px;
  display:grid; place-items:center;
  background: linear-gradient(135deg, rgba(95,155,140,0.32), rgba(160,195,130,0.22));
  border: 1px solid rgba(160,195,130,0.28);
}

.figure-strip{
  margin-top: 18px;
  display:grid;
  grid-template-columns: repeat(4, minmax(0,1fr));
  gap: 14px;
}
.figure{
  margin:0;
  background: rgba(255,255,255,0.05);
  border: 1px solid rgba(255,255,255,0.14);
  border-radius: var(--radius);
  overflow:hidden;
}
.figure img{
  width:100%;
  height: 230px;
  object-fit: cover;
  object-position: 50% 5%;
  object-position: 50% 0%;
  display:block;
  background: rgba(255,255,255,0.04);
}
.figure figcaption{
  padding: 10px 12px;
  color: var(--muted);
  font-size: 0.92rem;
  line-height: 1.5;
}

.filters{
  display:flex;
  flex-wrap:wrap;
  gap:10px;
  margin-bottom: 14px;
  padding-left: 4px;
}
.pill{
  background: var(--panel2);
  border: 1px solid var(--border);
  color: var(--muted);
  padding: 10px 12px;
  border-radius: 999px;
  cursor:pointer;
  font-weight: 650;
}
.pill.is-active{
  background: rgba(250,200,70,0.16);
  border-color: rgba(250,200,70,0.30);
  color: var(--text);
}

.pubs{
  display:grid;
  gap: 12px;
}
.pub{
  background: var(--panel);
  border: 1px solid var(--border);
  border-radius: var(--radius);
  padding: 14px 16px;
}
.pub h3{margin: 0 0 8px 0; font-size: 1.05rem}
.pub-meta{margin:0 0 8px 0; color: var(--muted)}
.pub-links a{
  color: var(--c4);
  text-decoration:none;
  font-weight: 700;
}
.pub-links a:hover{text-decoration:underline}

.callout{
  margin-top: 16px;
  padding: 16px;
  border-radius: var(--radius);
  border: 1px solid rgba(95,155,140,0.32);
  background: linear-gradient(135deg, rgba(95,155,140,0.18), rgba(35,60,75,0.20));
}
.callout h3{margin: 0 0 8px 0}
.callout p{margin:0; color: var(--muted); line-height: 1.6}

.note{
  margin-top: 14px;
  color: var(--muted);
}

.contact-card{
  display:grid;
  grid-template-columns: repeat(3, minmax(0,1fr));
  gap: 14px;
  padding: 16px;
  border-radius: var(--radius);
  border: 1px solid var(--border);
  background: var(--panel2);
}
.contact-card h3{margin: 0 0 8px 0}
.contact-card p{margin:0; color: var(--muted); line-height:1.6}
.contact-card a{color: var(--c3); text-decoration:none; font-weight: 700}
.contact-card a:hover{text-decoration:underline}
.sep{color: rgba(255,255,255,0.35); margin: 0 6px}
.muted{color: var(--muted); font-weight: 500}

.footer{
  margin-top: 22px;
  padding-top: 14px;
  border-top: 1px solid var(--border);
  color: rgba(255,255,255,0.62);
}

/* Screen reader only */
.sr-only{
  position:absolute !important;
  width:1px; height:1px;
  padding:0; margin:-1px;
  overflow:hidden; clip:rect(0,0,0,0);
  white-space:nowrap; border:0;
}

/* Responsive */
@media (max-width: 1100px){
  .figure-strip{grid-template-columns: repeat(2, minmax(0,1fr))}
}
@media (max-width: 980px){
  .hero-inner{grid-template-columns: 1fr}
  .cards-4{grid-template-columns: repeat(2, minmax(0,1fr))}
  .figure-strip{grid-template-columns: 1fr}
  .contact-card{grid-template-columns: 1fr}
}
@media (max-width: 760px){
  .nav-toggle{display:inline-flex}
  .nav-links{
    display:none;
    position:absolute;
    right: 20px;
    top: 64px;
    width: min(360px, calc(100% - 40px));
    padding: 12px;
    border-radius: var(--radius);
    background: rgba(15,29,37,0.92);
    border: 1px solid var(--border);
    box-shadow: var(--shadow-soft);
    backdrop-filter: blur(10px);
    flex-direction: column;
    align-items: stretch;
    gap: 8px;
    z-index: 50;
  }
  .nav-links a{padding: 12px 12px}
  .nav-links.is-open{display:flex}
  .two-cols{grid-template-columns: 1fr}
}



/* Portrait */
.portrait{
  width:100%;
  height: auto;
  max-height: 520px;
  object-fit: contain;
  border-radius: 16px;
  border: 1px solid var(--border);
  margin-bottom: 12px;
  background: rgba(255,255,255,0.04);
}
@media (max-width: 720px){
  .portrait{ max-height: 420px; }
}

/* Tabs */
.tabs{
  display:flex;
  flex-wrap:wrap;
  gap:10px;
  margin: 14px 0 12px;
}
.tab-btn{
  background: var(--panel2);
  border: 1px solid var(--border);
  color: var(--muted);
  padding: 10px 12px;
  border-radius: 999px;
  cursor:pointer;
  font-weight: 700;
}
.tab-btn.is-active{
  background: rgba(160,195,130,0.18);
  border-color: rgba(160,195,130,0.35);
  color: var(--text);
}
.tab-panels{
  margin-top: 10px;
}
.tab-panel{display:none}
.tab-panel.is-active{display:block}

/* Timeline */
.timeline{
  display:grid;
  gap: 12px;
}
.timeline-item{
  display:grid;
  grid-template-columns: 160px 1fr;
  gap: 14px;
  padding: 14px 14px;
  border-radius: var(--radius);
  border: 1px solid var(--border);
  background: var(--panel2);
}
.timeline-date{
  color: var(--muted);
  font-weight: 800;
}
.timeline-body p{
  margin: 8px 0 0 0;
  color: var(--muted);
  line-height: 1.6;
}
.inst{
  display:flex;
  align-items:center;
  gap: 10px;
}
.inst-logo{
  width: 40px;
  height: 40px;
  border-radius: 14px;
  display:grid;
  place-items:center;
  background: linear-gradient(135deg, rgba(255,125,45,0.35), rgba(250,200,70,0.30));
  border: 1px solid rgba(255,255,255,0.14);
  font-weight: 900;
  color: rgba(255,255,255,0.92);
  letter-spacing: 0.04em;
}
.inst-name{
  font-weight: 800;
}

/* Clean lists */
.clean-list{
  margin: 0;
  padding-left: 18px;
  color: var(--muted);
  line-height: 1.65;
}
.clean-list li{
  margin: 10px 0;
}
.clean-list.numbered{
  padding-left: 22px;
}
.clean-list .year{
  display:inline-block;
  min-width: 72px;
  color: rgba(255,255,255,0.88);
  font-weight: 800;
}

/* Better link styling inside long lists */
.clean-list a{
  color: var(--c4);
  text-decoration: none;
  font-weight: 700;
}
.clean-list a:hover{
  text-decoration: underline;
}

/* Responsive timeline */
@media (max-width: 760px){
  .timeline-item{
    grid-template-columns: 1fr;
  }
}


/* Technical decoration */
.hero, .section{position:relative; overflow:hidden;}
.decor{
  position:absolute;
  pointer-events:none;
  user-select:none;
  opacity:0.16;
  filter: drop-shadow(0 12px 30px rgba(0,0,0,0.35));
  z-index: 1;
}

.decor-hero-wallpaper{
  top: 96px;
  right: -220px;
  width: 980px;
  max-width: 72vw;
  opacity: 0.20;
  transform: rotate(-2deg);
}

.decor-hero-gn01{
  top: -70px;
  left: -220px;
  width: 520px;
  max-width: 44vw;
  opacity: 0.14;
  transform: rotate(10deg);
}

.decor-research-a3f{
  top: 10px;
  right: -260px;
  width: 760px;
  max-width: 60vw;
  opacity: 0.12;
  transform: rotate(3deg);
}

.decor-outputs-gnomad{
  top: -120px;
  left: -260px;
  width: 980px;
  max-width: 70vw;
  opacity: 0.10;
  transform: rotate(-4deg);
}

.decor-cv-gnomad{
  bottom: -140px;
  right: -260px;
  width: 980px;
  max-width: 70vw;
  opacity: 0.08;
  transform: rotate(5deg);
}

@media (max-width: 760px){
  .decor{display:none;}
}

.section .container{position:relative; z-index:2;}


/* ---- Improvements (2026-01) ---- */
.site-header{
  position: sticky;
  top: 0;
  z-index: 50;
}
.site-header.is-sticky{
  background: rgba(255,255,255,0.78);
  backdrop-filter: blur(10px);
  border-bottom: 1px solid rgba(35,60,75,0.10);
}
.site-header.is-sticky .nav{
  padding: 10px 0;
}

.hero-tagline{
  margin: 10px 0 14px;
  max-width: 70ch;
  font-weight: 850;
  letter-spacing: 0.2px;
  color: var(--c1);
  text-shadow: 0 1px 0 rgba(255,255,255,0.55);
}

.pub-controls{
  display:flex;
  flex-wrap:wrap;
  gap: 12px;
  align-items:center;
  justify-content:space-between;
  margin-top: 12px;
}
.pub-tools{
  display:flex;
  flex-wrap:wrap;
  gap: 10px;
  align-items:center;
}
.input, .select{
  background: var(--panel2);
  border: 1px solid var(--border);
  color: var(--text);
  padding: 10px 12px;
  border-radius: 999px;
}
.input{ min-width: 240px; }
.select{ min-width: 190px; }

.pub-count{
  margin: 10px 0 14px;
  color: var(--muted);
}

.pub-top{
  display:flex;
  align-items:flex-start;
  justify-content:space-between;
  gap: 14px;
}
.pub-actions{
  display:flex;
  gap: 8px;
  align-items:center;
  flex-shrink: 0;
}
.pub-link{
  display:inline-flex;
  align-items:center;
  gap: 6px;
  text-decoration:none;
  padding: 6px 10px;
  border-radius: 999px;
  border: 1px solid rgba(250,200,70,0.30);
  background: rgba(250,200,70,0.10);
  color: var(--text);
  font-weight: 700;
  font-size: 0.85rem;
}
.pub-link:hover{ filter: brightness(1.06); }

.tags{
  display:flex;
  flex-wrap:wrap;
  gap: 8px;
  margin-top: 8px;
}
.tag{
  font-size: 0.78rem;
  padding: 6px 10px;
  border-radius: 999px;
  border: 1px solid var(--border);
  background: rgba(255,255,255,0.04);
  color: var(--muted);
  font-weight: 650;
}

.featured-block{
  margin: 18px 0 14px;
  padding: 14px 14px 16px;
  border-radius: calc(var(--radius) + 6px);
  border: 1px solid var(--border);
  background: linear-gradient(135deg, rgba(95,155,140,0.10), rgba(35,60,75,0.10));
}
.featured-head{
  display:flex;
  flex-wrap:wrap;
  gap: 10px;
  align-items:baseline;
  justify-content:space-between;
  margin-bottom: 10px;
}
.featured-head h3{ margin: 0; }
.featured-head .muted{ margin: 0; color: var(--muted); max-width: 70ch; }

.featured-pubs{
  display:grid;
  gap: 12px;
  grid-template-columns: repeat(auto-fit, minmax(220px, 1fr));
}
.pub--featured h3{ font-size: 1.0rem; }
.pub--featured .pub-meta{ font-size: 0.92rem; }
.pub-year{ margin: 0 0 6px 0; color: var(--muted); font-weight: 650; }

a:focus-visible,
button:focus-visible,
input:focus-visible,
select:focus-visible{
  outline: 3px solid rgba(250,200,70,0.55);
  outline-offset: 3px;
}

@media (prefers-reduced-motion: reduce){
  *{ scroll-behavior: auto !important; }
}